Jonathan grew up in Edinburgh, attending Heriot-Watt University where he completed a BSc in Chemistry with Pharmaceutical Chemistry, working on the synthesis of Botulinum B and Tetanus toxin light chains for use in super resolution microscopy in the Rickman Lab of the Institute of Biological Chemistry, Biophysics and Bioengineering (IB3). He then attended the University of Edinburgh, where he did an MSc in Synthetic Biology & Biotechnology and developed, as part of his dissertation, a toolkit for the high-throughout, in vivo directed evolution of enzymes on the Taxol biosynthetic pathway in the Rios Lab of the Institute of Bioengineering (IBioE). Whilst completing his MSc, he received the MacQuarrie Fellowship, allowing him to work as Scientist-in-Residence at George Watson’s College, in particular arranging and hosting lab placements for school pupils. Right after this, he began his PhD in Edinburgh’s Institute of Quantitative Biology, Biochemistry and Biotechnology (IQB3), under the supervision of Dr Stephen Wallace and Prof Dominic Campopiano. For his PhD project, he will be working on metabolic engineering approaches to the diastereoselective synthesis of a class of natural products with pharmaceutical applications.